Which one of the following sentences demonstrates correct comma usage? "By examining the state of Bhutan’s natural resources the pressures on ecosystems and different management responses, the domain of ecological diversity and resilience is intended to describe the impact of domestic supply and demand on Bhutan’s ecosystems." "By examining the state of Bhutan’s natural resources, the pressures on ecosystems, and different management responses the domain of ecological diversity and resilience is intended to describe the impact of domestic supply and demand on Bhutan’s ecosystems." "By examining the state of Bhutan’s natural resources, the pressures on ecosystems, and different management responses, the domain of ecological diversity and resilience is intended to describe the impact of domestic supply and demand on Bhutan’s ecosystems." "By examining the state of Bhutan’s natural resources the pressures on ecosystems, and different management responses, the domain of ecological diversity and resilience is intended to describe the impact of domestic supply, and demand on Bhutan’s ecosystems."